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I use several different programs to access the same file in the Dropbox on Windows, iOS and OS X. All those different apps/programs update this file. After some days/weeks the link I use in the iOS App to the file changes and the App does not work anymore with this file. When, under which circumstances does Dropbox changes the link of a file?

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You mean the public url of the file? – Bibhas May 11 '12 at 17:09
Yey, the public URL. – powtac May 14 '12 at 12:41

Whenever you change the path of the file on the device you are syncing from,
the public URL changes to reflect the change in path.

[I can only confirm this for Windows based devices.]

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