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This has been happening a lot lately, mostly on YouTube. I don't know if it's YouTube, Adobe Flash or Firefox:
This will happen even half-way through playing a video, it will randomly turn into this image. I have to restart Firefox to continue to play YouTube videos.
It is accused of being a bug (of certain versions of Flash Player) and an out of memory notification.
Adobe Forums: Gray circle with white exclamation mark on white background
It's your Adobe Flash.
Try restarting your browser. Then go back to youtube.com and try to update your Java/Adobe/default Flash program.
If it's happening across platforms and regardles of video accelerator than it is clearly a YouTube problem and not specific to one's PC or OS.
I think it's Firefox. This happened to me on Google Chrome, and the grey excalimation mark happened on YouTube, and another site I was watching an anime on. I tried YouTube on Internet Explorer, and it works fine.
I suggest uninstalling and reinstalling Firefox.
Most likely it's Adobe Flash as others have stated. If restarting doesn't help I suggest switching to HTML5 on YouTube. You can do this by following the following link: http://www.youtube.com/html5
Good Luck!
